The order Dipsacales is classified in Euasterids II and comprises six families following APG: Caprifoliaceae, Diervillaceae, Linnaeaceae, Dipsacaceae, Valerianaceae and Morinaceae. Despite its relatively small size, many phylogenetic questions on intergeneric and interfamilial relations still remain in this order. New sequences of both 5 and 3 region of the plastid ndhF gene are generated and combined with published ndhF sequence data; a combination with available rbcL sequences and a morphological data set is also performed. All major lineages within Dipsacales are represented. Parsimony analysis based on ndhF data shows Heptacodium as sister to all members of Caprifoliaceae; combined algorithms suggest an unresolved basal position in the Dipsacales. Trees inferred from all data sets indicate a paraphyletic nature of the tribe Patrinieae and suggest that Nardostachys can no longer be regarded as the basalmost member of Valerianaceae. Morinaceae are strongly monophyletic and behave as a basal clade to the Valerianaceae-Dipsacaceae clade. In classifications of Dipsacales the genus Triplostegia is treated as a member of Valerianaceae, Dipsacaceae, or even as a monogeneric family. The results presented, indicate that the position of Triplostegia remains equivocal: molecular data suggest a basal position in Dipsacaceae, while the morphological matrix and the total combined data set show Triplostegia as sister to all Valerianaceae.

Key words: combined analyses, Dipsacales, morphology, ndhF, phylogeny, rbcL